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School of Forest, Fisheries, & Geomatics Sciences

School of Forest, Fisheries, & Geomatics Sciences

Timothy A. Martin

Professor, Tree Physiology

Tim Martin joined the School in 1999 with teaching and research responsibilities in Tree Physiology. From 1997-1999, he worked as a Research Associate with the Forest Biology Research Cooperative; he now serves as a Co-Director for the FBRC.

His research program focuses on the physiological mechanisms controlling productivity, carbon and water flux in trees and forests. He teaches undergraduate and graduate courses in tree physiology and forest management.

Courses Taught

IdentifierCourse Name
FOR3200C  Foundations in Natural Resources and Conservation 
FOR3342C  Tree Biology 
FNR4623C  Integrated Natural Resource Management 
FOR6340  Physiology of Forest Trees 


359 Newins-Ziegler Hall, PO Box 110410
Gainesville, FL 32611-0410
(352) 846-0866

  • Education

    • PhD, Ecosystem Analysis, University of Washington, 1997
    • MS, Tree Physiology, University of Georgia, 1992
    • BS, Forest Management, Oklahoma State University, 1990

  • Publications

    • Ribeiro, C.L., D. Conde, K.M. Balmant, C. Dervinis, M.G.Johnson, A.P.MGrath, P. Szewczyk, F. Unda, C.A. Finegan, H.W.Schmidt, B. Miles, D.R. Drost, E. Novaes, C.A. Gonzalez-Benecke, G.F. Peter, J.G. Burleigh, T.A. Martin, S.D. Mansfield, G. Chang, N.J. Wickett, and M. Kirst. 2020. The uncharacterized gene EVE contributes to vessel element dimensions in Populus. Proceedings of the National Academy of Sciences 117:5059-5066.
    • Yasin, G., M.H. Nawaz, T.A. Martin, N.K. Niazi, S. Gul, M.T.B. Yousaf. 2019. Evaluation of agroforestry carbon storage status and potential in irrigated plains of Pakistan. Forests 10:640
    • Bracho, R., J.G. Vogel, R.E. Will, A. Noormets, L.J. Samuelson, E.J. Jokela, C.A. Gonzalez-Benecke, S.A. Gezan, D. Markewitz, J.R. Seiler, B.D. Strahm, R.O. Teskey, T.R. Fox, M.B. Kane, M.A. Laviner, K.M. McElligot, J. Yang, W. Lin, C.R. Meek, J. Cucinella, M.K. Akers, and T.A. Martin. 2018. Carbon accumulation in loblolly pine plantations is increased by fertilization across a soil moisture availability gradient. Forest Ecology and Management 424:39-52.
    • Martin, T.A., D.C. Adams, M.J. Cohen, R.M. Crandall, C.A. Gonzalez-Benecke, J.A. Smith, and J.G. Vogel. 2017. Managing Florida’s plantation forests in a changing climate. In: Florida's Climate: Changes, Variations, and Impacts, E.P. Chassignet, J.M. Jones, V. Misra, and J. Obeysekera, Eds. Florida Climate Institute, Gainesville, Florida. pp. 269-295.
    • Eigenbrode, S.D., T.A. Martin, L. Wright Morton, J. Colletti, P. Goodwin, R. Gustafson, D. Hawthorne, A. Johnson, L. Mercado, S. Pearl, T. Richard, and M. Wolcott. 2017. Leading large transdisciplinary projects addressing social-ecological systems: A primer for project directors. USDA National Institute for Food and Agriculture, Washington, D.C. 69 p.
    • Quesada,T., L.M. Parisi, D.A. Huber, S.A. Gezan, T.A. Martin, J.M. Davis, and G.F. Peter. 2017. Genetic control of growth and shoot phenology in juvenile loblolly pine (Pinus taeda L.) clonal trials. Tree Genetics and Genomes 13(3):1-15.
    • Gonzalez-Benecke, C.A., R.O. Teskey, H. Dinon Aldridge, and T.A. Martin. 2017. Pinus taeda forest growth predictions in the 21st century vary with site mean annual temperature and site quality. Global Change Biology
    • Samuelson, L.J., T.A. Stokes, J.R. Butnor, K.H. Johnsen, C.A. Gonzalez-Benecke, T.A. Martin, W. P. Cropper, Jr., P.H. Anderson, M.R. Ramirez, and J.C. Lewis. 2017. Ecosystem carbon density and allocation across a chronosequence of longleaf pine forests. Ecological Applications 27:244-259.
    • Monroe, M.C., J.J.T. Ireland, and T.A. Martin. 2015. Integration of forestry research and extension in an online graduate course. Journal of Forestry 113:240-247.
    • Will, R.E., T.R. Fox, M.K. Akers, J.-C. Domec, C.A. Gonzalez-Benecke, E.J. Jokela, M. Kane, M.A. Laviner, G. Lokuta, D. Markewitz, M.A. McGuire, C. Meek, A. Noormets, L.J. Samuelson, J.R. Seiler, B. Strahm, R.O. Teskey, J.G. Vogel, E.J. Ward, J. West, D.S. Wilson, T.A. Martin. 2015. A range-wide experiment to investigate nutrient and soil moisture interactions in loblolly pine plantations. Forests 6:2014-2028.
    • Eigenbrode, S.D., L. Wright Morton, and T.A. Martin. 2014. Big interdisciplinarity to address climate change and agriculture: Lessons from three USDA Coordinated Agricultural Projects. Journal of Soil and Water Conservation 69:170A-175A.
    • Resende, M.F.R., Jr., P. Munoz, M.D.V. Resende, D.J. Garrick, R.L. Fernando, J.M. Davis, E.J. Jokela, T.A. Martin, G.F. Peter, and M. Kirst. 2012. Accuracy of genomic selection methods in a standard dataset of loblolly pine (Pinus taeda L.). Genetics 190:1503-1510.
    • Bracho, R., G. Starr, H.L. Gholz, T.A. Martin, W.P. Cropper, Jr., and H.W. Loescher. 2012. Controls on carbon dynamics by ecosystem structure and climate for southeastern U.S. slash pine plantations. Ecological Monographs 82:101-128.
    • Gonzalez-Benecke, C.A., T.A. Martin, A. Clark, III, and G.F. Peter. 2010. Water availability and genetic effects on wood properties of loblolly pine (Pinus taeda L.). Canadian Journal of Forest Research 40:2265-2277.
    • Staudhammer, C.L., E.J. Jokela, and T.A. Martin. 2009. Competition dynamics in pure- versus mixed-family stands of loblolly and slash pine in the southeastern United States. Canadian Journal of Forest Research 39: 396-409.
    • Powell, T.L., H.L. Gholz, K.L. Clark, G. Starr, W.P. Cropper, Jr., and T.A. Martin. 2008. Carbon exchange of a mature, naturally-regenerated pine forest in north Florida. Global Change Biology 14: 2523-2538.
    • Baltunis, B.S., T.A. Martin, D.A. Huber, and J.M. Davis. 2008. Inheritance of foliar stable carbon isotope discrimination and third-year height in Pinus taeda clones on contrasting sites in Florida and Georgia. Tree Genetics and Genomes 4: 797-807.
    • Emhart, V.I., T.A. Martin, T.L. White, and D.A. Huber. 2006. Genetic variation in basal area increment phenology and its correlation with growth rate in loblolly and slash pine families and clones. Canadian Journal of Forest Research 36: 961-971.
    • Martin, T.A., P.M. Dougherty, M.A. Topa and S.E. McKeand. 2005. Strategies and case studies for incorporating ecophysiology into southern pine tree improvement programs. Southern Journal of Applied Forestry 29: 70-79.
    • Martin, T.A., and E.J. Jokela. 2004. Developmental patterns and nutrition impact radiation use efficiency components in southern pine stands. Ecological Applications 14: 1839-1854.
    • McGarvey, R.C., T.A. Martin, and T.L. White. 2004. Integrating within-crown variation in net photosynthesis in loblolly and slash pine families. Tree Physiology 24: 1209-1220.
    • Martin, T.A., K.J. Brown, T.M. Hinckley, J. Kučera, F.C. Meinzer and D.G. Sprugel. 2001. Control of transpiration in a 220-year-old Abies amabilis forest. Forest Ecology and Management 152: 211-224.
    • Martin, T.A., K.H. Johnsen, and T.L. White. 2001. Ideotype development in southern pines: Rationale and strategies for overcoming scale-related obstacles. Forest Science 47: 21-28.
    • Martin, T.A., K.J. Brown, J. Cermák, R. Ceulemans, J. Kucera, F.C. Meinzer, J.S. Rombold, D.G. Sprugel and T.M. Hinckley. 1997. Crown conductance and tree and stand transpiration in a second-growth Abies amabilis forest. Canadian Journal of Forest Research 27: 797-808.
    • Martin, T.A., R.O. Teskey and P.M. Dougherty. 1994. Movement of respiratory CO2 in stems of loblolly pine (Pinus taeda L.) seedlings. Tree Physiology 14: 481-495.